Flora and Funga records
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Flora and Funga records
Notulae to the Italian native vascular flora: 21
expand article infoFabrizio Bartolucci‡, Gianniantonio Domina§, Michele Adorni|, Alessandro Alessandrini, Nicola M. G. Ardenghi#, Carlo Argenti¤, Enrico Banfi«, Gianmaria Bonari»˄, Emanuel Bonivento˅, Diego Cantini», Pasquale Cerbone¦, Maria Carla de Francescoˀ, Riccardo De Vivoˁ, Emanuele Del Guacchio₵, Adriano Di Pietroℓ, Romeo Di Pietro₰, Luigi Forte₱, Gabriele Galasso«, Roberto Gennaio₳, Luigi Ghillani₴, Günter Gottschlich₣, Michele Lonati₮, Quintino Giovanni Manni₦, Francesca Mantino₱, Carlo Mazzera₭, Francesco Minutillo₲, Ginevra Nota₮, Giuseppe Pappagallo‽, Elisa Proietti₰, Leonardo Rosati₩, Giovanni Salerno₸, Lucia Antonietta Santoianniˀ, Alberto Selvaggi‡‡, Giancarlo Tondi§§, Thomas Wilhalm||, Corrado Zanni¶¶, Lorenzo Lastrucci##
‡ Dipartimento di Biologia Ambientale, Sapienza Università di Roma, Roma, Italy
§ Dipartimento di Scienze Agrarie, Alimentari e Forestali (SAAF), Università di Palermo, Palermo, Italy
| Orto Botanico, Sistema Museale di Ateneo, Università di Parma, Parma, Italy
¶ Via G. Pilati, Bologna, Italy
# Orto Botanico, Sistema Museale di Ateneo, Università di Pavia, Pavia, Italy
¤ Via Pietriboni, Belluno, Italy
« Sezione di Botanica, Museo di Storia Naturale di Milano, Milano, Italy
» Dipartimento di Scienze della Vita, Università degli studi di Siena, Siena, Italy
˄ National Biodiversity Future Center, Palermo, Italy
˅ Via Cavour, Biella, Italy
¦ Via Vittorio Imbriani, Napoli, Italy
ˀ Dipartimento di Bioscienze e Territorio, Università del Molise, Termoli, Italy
ˁ Via XX Settembre, Parma, Italy
₵ Dipartimento di Biologia, Università di Napoli Federico II, Napoli, Italy
ℓ Via Boccea, Roma, Italy
₰ Dipartimento di Pianificazione, Design, Tecnologia dell’Architettura (PDTA), Sapienza Università di Roma, Roma, Italy
₱ Dipartimento di Bioscienze, Biotecnologie e Ambiente – Museo Orto Botanico, Campus Universitario “E. Quagliariello”, Università degli Studi di Bari “Aldo Moro”, Bari, Italy
₳ Vincenzo Bellini, Lecce, Italy
₴ Via Casalegno, Parma, Italy
₣ Hermann-Kurz-Str., Tübingen, Germany
₮ Dipartimento di Scienze Agrarie, Forestali e Alimentari, Università degli Studi di Torino, Grugliasco, Italy
₦ Via San Michele, Lecce, Italy
₭ Via Lituania, Parma, Italy
₲ Via Cuostile, Gaeta, Italy
‽ Istituto di Ricerca sulle Acque (CNR-IRSA), Bari, Italy
₩ Dipartimento di Scienze della Salute, Università degli Studi della Basilicata, Potenza, Italy
₸ Via O. Coccanari, Tivoli, Italy
‡‡ FHP - Flora Habitat Piemonte, Cuneo, Italy
§§ Via F. D’Ovidio, Roma, Italy
|| Museo di Scienze Naturali dell’Alto Adige, Bolzano, Italy
¶¶ Via Carlo Casalegno, Parma, Italy
## Sistema Museale di Ateneo, Università di Firenze, Firenze, Italy

Corresponding author: Fabrizio Bartolucci ( fabrizio.bartolucci@uniroma1.it )

Academic editor: Stefania Biondi
© 2026 Fabrizio Bartolucci, Gianniantonio Domina, Michele Adorni, Alessandro Alessandrini, Nicola M. G. Ardenghi, Carlo Argenti, Enrico Banfi, Gianmaria Bonari, Emanuel Bonivento, Diego Cantini, Pasquale Cerbone, Maria Carla de Francesco, Riccardo De Vivo, Emanuele Del Guacchio, Adriano Di Pietro, Romeo Di Pietro, Luigi Forte, Gabriele Galasso, Roberto Gennaio, Luigi Ghillani, Günter Gottschlich, Michele Lonati, Quintino Giovanni Manni, Francesca Mantino, Carlo Mazzera, Francesco Minutillo, Ginevra Nota, Giuseppe Pappagallo, Elisa Proietti, Leonardo Rosati, Giovanni Salerno, Lucia Antonietta Santoianni, Alberto Selvaggi, Giancarlo Tondi, Thomas Wilhalm, Corrado Zanni, Lorenzo Lastrucci.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation: Bartolucci F, Domina G, Adorni M, Alessandrini A, Ardenghi NMG, Argenti C, Banfi E, Bonari G, Bonivento E, Cantini D, Cerbone P, de Francesco MC, De Vivo R, Del Guacchio E, Di Pietro A, Di Pietro R, Forte L, Galasso G, Gennaio R, Ghillani L, Gottschlich G, Lonati M, Manni QG, Mantino F, Mazzera C, Minutillo F, Nota G, Pappagallo G, Proietti E, Rosati L, Salerno G, Santoianni LA, Selvaggi A, Tondi G, Wilhalm T, Zanni C, Lastrucci L (2026) Notulae to the Italian native vascular flora: 21. Italian Botanist 21: 237-253. https://doi.org/10.3897/italianbotanist.21.203684
Open Access

Abstract

In this contribution, new data concerning the distribution of native vascular flora in Italy are presented. It includes new records, confirmations, and exclusions to the Italian administrative regions for taxa in the genera Angelica, Cuscuta, Dianthus, Ficus, Hieracium, Lamium, Laurus, Lomelosia, Lophiolepis, Nerium, Orchis, Pimpinella, Potentilla, Senecio, Thalictrum, and Trifolium. New combinations in the genera Festuca (Festuca × stacei, Poaceae) and Anemone (Anemone alpina subsp. austroalpina, Ranunculaceae) are proposed. Nomenclatural and distribution updates, published elsewhere, and corrigenda are provided as Suppl. material 1.

Keywords

Endemic taxa, floristic data, Italy, nomenclature, taxonomy

How to contribute

The text for the new records, exclusions, and confirmations should be submitted electronically to Lorenzo Lastrucci (lorenzo.lastrucci@unifi.it). The corresponding specimen along with its scan or photograph have to be sent to FI Herbarium: Sezione di Botanica “Filippo Parlatore” del Museo di Storia Naturale, Via G. La Pira 4, 50121 Firenze (Italy). Those texts concerning nomenclatural novelties and typifications (only for accepted names) should be submitted electronically to: Fabrizio Bartolucci (fabrizio.bartolucci@gmail.com). Each text should be within 1,000 characters (spaces included).

Floristic records of native taxa

Angelica sylvestris L. subsp. sylvestris (Apiaceae)

+ PUG: Faeto (Foggia), Torrente Celone-Le Cesi, Monti Dauni (WGS84: 41.332769°N, 15.167281°E), bordure a megaforbie igrofile ai margini del torrente, 660 m, 21 September 2025, G. Pappagallo (FI). – Species confirmed for the flora of Puglia.

G. Pappagallo

Cuscuta palaestina Boiss. subsp. palaestina (Convolvulaceae)

+ TOS: Grosseto, oliveto abbandonato su suolo calcareo con vegetazione della classe Stipo-Trachynietea distachyae, a Poggio Bernarda, Parco Regionale della Maremma, presso Alberese (WGS84: 42.661889°N, 11.108611°E), 30 m a.s.l., 14 May 2025, G. Bonari, D. Cantini, V. González-García, G. Terranova (SIENA). – Species new to the flora of Toscana.

G. Bonari, D. Cantini

Dianthus rupicola Biv. subsp. rupicola (Caryophyllaceae)

+ LAZ: Ponza (Isole Ponziane, Latina) (WGS84: 40.917880°N, 12.958596°E), rupi, suolo vulcanico, 40 m, 11 September 2025, F. Minutillo (FI; Herb. Tondi). – Species new for the flora of Lazio.

F. Minutillo

Hieracium caesium (Fr.) Fr. subsp. brennerianum (Zahn) Gottschl. (Asteraceae)

+ VEN: Lozzo di Cadore (Belluno), Fienili Vialona, Val Longerin (WGS84: 46.490433°N, 12.412742°E), prato riva, 1075 m, 9 July 2025, leg. C. Argenti, det. G. Gottschlich (FI, Herb. Argenti); Danta di Cadore (Belluno), Ponte Mauria (WGS84: 46.560559°N, 12.499791°E), scarpata erbosa, 1300 m, 12 June 2025, leg. C. Argenti, det. G. Gottschlich (FI, Herb. Argenti). – Subspecies new for the flora of Veneto.

G. Gottschlich, C. Argenti

Hieracium cydoniifolium Vill. subsp. trichanthodium Zahn (Asteraceae)

+ ITALIA (TAA): Bozen, Vinschgau, Martell, 0,44 km S(S)W Schluderalm (WGS84: 46.52999°N, 10.7294°E), aufgelichteter subalpiner Nadelwald, Marteller Granit, 2063 m, 30 August 2024, leg. E. Spögler, Th. Wilhalm et al., rev. G. Gottschlich (FI, BOZ, Hb. Gottschlich-85396). – Subspecies new for the flora of Italy (Trentino-Alto Adige).

The subspecies is very similar to Hieracium prenanthoides, but is much more hairy on the peduncles than all subspecies of H. prenanthoides and thus represents a first transition to this species. So far it is only known from Switzerland (Valais, Bern, Grisons).

G. Gottschlich, T. Wilhalm

Hieracium dentatum Hoppe subsp. pseudoplantagineum Zahn (Asteraceae)

+ VEN: Rocca Pietore (Belluno), tra Passo Fedaia e Passo Padon, M.te Padon (WGS84: 46.459396°N, 11.890831°E), pascolo rupestre su silice, 2200 m, 20 July 2025, leg. C. Argenti, det. G. Gottschlich (FI, Herb. Argenti). – Subspecies new for the flora of Veneto.

G. Gottschlich, C. Argenti

Hieracium diaphanoides Lindeb. subsp. diaphanoides (Asteraceae)

+ VEN: Comelico Superiore (Belluno), Malga Coltrondo sul Col Quaternà (WGS84: 46.663205°N, 12.451475°E), margine bosco, 1880 m, 18 July 2025, leg. C. Argenti, det. G. Gottschlich (FI). – Subspecies new for the flora of Veneto.

G. Gottschlich, C. Argenti

Hieracium froelichianum H.Buek subsp. subvulsum (Zahn) Gottschl. & Greuter (Asteraceae)

+ VEN: Livinallongo del Col di Lana (Belluno), Salere sul versante nord del M.te Padon (WGS84: 46.485279°N, 11.847825°E), rupestre su silice, 1950 m, 2 August 2023, leg. C. Argenti, det. G. Gottschlich (Herb. Argenti); Rocca Pietore (Belluno), tra Passo Fedaia e Passo Padon, M.te Padon (WGS84: 46.454776°N, 11.888661°E), pascolo rupestre su silice, 2100 m, 20 July 2025, leg. C. Argenti, det. G. Gottschlich (FI). – Subspecies new for the flora of Veneto.

G. Gottschlich, C. Argenti

Hieracium levicaule Jord. subsp. pseudosenile (Zahn) Zahn (Asteraceae)

+ VEN: Comelico Superiore (Belluno), riva dx T. Padola a Padola (WGS84: 46.610186°N, 12.478050°E), margine bosco, 1200 m, 18 June 2025, leg. C. Argenti, det. G. Gottschlich (FI, Herb. Argenti). – Subspecies new for the flora of Veneto.

G. Gottschlich, C. Argenti

Hieracium pallescens Waldst. & Kit. subsp. ovale (Zahn) Gottschl. (Asteraceae)

+ VEN: Feltre (Belluno), Collesei di Anzù (WGS84: 46.007972°N, 11.946841°E), rupestre su calcare, 250 m, 26 April 2025, leg. C. Argenti, det. G. Gottschlich (FI, Herb. Argenti). – Subspecies new for the flora of Veneto.

G. Gottschlich, C. Argenti

Lamium album L. subsp. album (Lamiaceae)

+ LAZ: Accumoli (Rieti), M. L’Inversaturo (WGS84: 42.691957°N, 13.303831°E), radura in faggeta; suolo marnoso-arenaceo, 1500 m, 7 June 1992, G. Tondi (FI); Amatrice (Rieti), Fosso della Pacina Monte Pelone (Monti della Laga) (WGS84: 42.63164°N, 13.38777°E), prateria di alte erbe e megaforbie, 30 July 2024, A. Di Pietro (HFLA). – Species new for the flora of Lazio.

Lamium album is regarded as wrongly reported for Lazio by Bartolucci et al. (2024). According to Mennema (1989) the collected specimen shows an oblique internal ring of hairs and light yellow pollen grains, both traits being considered diagnostic of L. album vs. L. maculatum L.

G. Tondi, A. Di Pietro

Lomelosia stellata (L.) Raf. (Caprifoliaceae)

- ITALIA (EMR). Species to be excluded from the flora of Italy (Emilia-Romagna).

Plants referred to this species were discovered along the Taro river in 2008 and recorded as new for the flora of Italy (Ghillani and Alessandrini 2012). The identification was confirmed by L. Villar (Spain) based on the analysis of photographic material. However, the correctness of the identification was questioned in the Acta Plantarum Forum by Valerio Lazzeri (https://www.actaplantarum.org/forum/viewtopic.php?p=826601), who proposed to refer the plant to Lomelosia micrantha. The analysis of fresh and herbarium specimens allowed us to exclude L. stellata and to attribute the taxon to the proposed species (see the following Notula).

L. Ghillani, M. Adorni, A. Alessandrini

Lomelosia micrantha (Desf.) Greuter & Burdet (Caprifoliaceae)

+ ITALIA (EMR): Medesano (Parma), fiume Taro presso Felegara (WGS84: 44.714190°N, 10.111376°E), prato arido su terrazzo alluvionale consolidato, 125 m, 3 June 2025, L. Ghillani, M. Adorni (FI). – Species new for the flora of Italy (Emilia-Romagna).

This species was identified using Flora Europaea (Jasiewicz 1976), under the name Scabiosa micrantha Desf., by analyzing photographic material and herbarium samples available online. The involucel-corona of Lomelosia micrantha is significantly narrower than that of Lomelosia stellata. Moreover, the involucral bracts are entire, while in L. stellata they are frequently divided (Devesa 2007). Lomelosia micrantha occurs from the central Balkan Peninsula to central Asia. This record represents the westernmost locality of the species’ range. The population of the Taro river consists of approximately 100 individuals and is the only one known for this species in Italy.

L. Ghillani, M. Adorni, A. Alessandrini

Lophiolepis morisiana (Rchb.f.) Del Guacchio, Bureš, Iamonico & P.Caputo (Asteraceae)

− CAM: Species to be excluded from the flora of Campania.

Lophiolepis morisiana is recorded in Campania only for the Partenio massif (“Monti di Avella”, Avellino) (Moraldo & La Valva, 1989, under the name Cirsium morisianum Rchb.f.). In fact, the locality “Lago Matese” (Caserta) reported by Bianchini (1988) after Martelli and Tanfani (1891) is erroneous, as those authors indicated “Cirsium eriophorum L.” (currently L. eriophora (L.) Del Guacchio, Bureš, Iamonico & P.Caputo), in turn to be interpreted sensu Arcangeli (1892, under Cnicus) (cf. Martelli and Tanfani 1891, p. 173) and likely to be referred to L. tenoreana (L.) (Petr.) Del Guacchio, Bureš, Iamonico & P.Caputo (Del Guacchio, pers. obs.). However, the indication by Moraldo and La Valva (1989) is not supported by any available specimen (NAP!), whereas in the same massif and surrounding areas, the similar L. lacaitae (L.) Del Guacchio, Bureš, Iamonico & P.Caputo, although not reported by them, indeed occurs (Cerbone and Del Guacchio, in prep.).

P. Cerbone, E. Del Guacchio

Orchis patens Desf. subsp . brevicornis (Viv.) K.Richt . (Orchidaceae)

+ EMR: Bardi (Parma), Passo del Pellizzone (WGS84: 44.67979°N, 09.74392°E), prato arido cespugliato, 1015 m, 26 May 2025, leg. C. Mazzera, det. C. Mazzera, M. Adorni (FI). – Subspecies new for the flora of Emilia-Romagna.

Orchis patens occurs only in northern Africa (Algerian-Tunisian Atlas) and north-western Italy, while Canarian populations were recently treated as O. canariensis Lindl. African populations are referred to the autonymic subspecies, while Italian ones, previously reported only for Liguria, are treated as O. patens subsp. brevicornis (Calevo et al. 2021, 2023). The population reported here consists of only nine individuals scattered over a surface of 15 m2 in a semi-natural dry grassland with southern aspect. It is the highest (the first one above 1,000 m a.s.l.) and the northernmost of the range of this species.

C. Mazzera, M. Adorni

Origanum onites L.(Lamiaceae)

+ PUG: Gallipoli (Lecce), Loc. Itri (WGS84: 40.007993°N, 18.035439°E), Cava per l’estrazione di conci di calcare “carparo” dismessa, 20 m s.l.m., 3 September 2025, leg. R. Gennaio, det. R. Gennaio, Q.G. Manni (FI). – Species new for the flora of Puglia.

R. Gennaio, Q.G. Manni

Pimpinella anisoides V.Brig. (Apiaceae)

− PUG: Species to be excluded from the flora of Puglia.

Pimpinella anisoides is reported as occurring in Puglia in the latest checklist of the vascular flora native to Italy (Bartolucci et al. 2024) based on Bianco et al. (1998) as well as in the recent Flora d’Italia (Pignatti et al. 2018). In Bianco et al. (1998), this species is reported only in the appendix among the sporadic species recorded in two vegetation relevés, and no deposited specimens in BI or CAT have been traced, thus raising doubts about the actual occurrence of the species in Puglia. Earlier, Scaramuzzi (1952) and Bianco (1962)— the latter confirming Scaramuzzi’s record and treating it as a new species for Puglia — reported the species from the Murgia di S. Elio (Altamura, Bari). We revised the specimen collected by Scaramuzzi and preserved in BI [Murgia di S. Elio, 18 June 1949, F. Scaramuzzi (BI No. 23744)] and we attribute it to Ammoides pusilla (Brot.) Breistr. We also revised another specimen preserved in BI under Pimpinella anisoides [Bosco “Selva” – Alberobello, 27 July 1952–27 August 1952, M. Iannice (BI No. 27610)], which we likewise attribute to Ammoides pusilla (Brot.) Breistr.

L. Forte, F. Mantino

Potentilla verna L. (Rosaceae)

+ LAZ: Amatrice (Rieti), Cardito, between Casa Cantoniera and Poggio Cancelli SS577 (WGS84: 42.582073°N, 13.319585°E), mesophilous grasslands, 1275 m a.s.l., 24 August 2024, leg. R. Di Pietro and P. Fortini, det. R. Di Pietro and E. Proietti (FI). – Species confirmed for flora of Lazio.

The species is not reported for the flora of Lazio either in Anzalone et al. (2010) or in Lucchese (2024). Tondi and Plini (1995) do not include P. verna among the species occurring in the Lazio slope of the Gran Sasso-Laga National Park, while Conti and Bartolucci (2016) indicate this species as having been erroneously included in the past for the flora of the Gran Sasso-Monti della Laga National Park.

R. Di Pietro, E. Proietti

Senecio apenninus Tausch (Asteraceae)

+ EMR: Monchio delle Corti (Parma), tra M. Paitino e M. Sillara (WGS84: 44.371618°N, 10.056568°E), formazioni erbose con affioramenti rocciosi, 1772 m, 15 July 2025, M. Adorni, R. De Vivo (FI); Monchio delle Corti (Parma), tra Cima Canuti e M. Pitturina (WGS84: 44.351209°N, 10.110477°E), praterie di cresta, 1735 m, 8 July 2023 (Herb. M. Adorni). – Species new for the flora of Emilia-Romagna.

This species has recently been re-evaluated by Conti et al. (2019), who consider it endemic to the central Apennines, where it is widespread in Marche, Umbria, Lazio, Abruzzo, and doubtfully occurring in Molise. The discovery in the Tuscan-Emilian Apennines significantly shifts the limit of its range northward.

M. Adorni, R. De Vivo

Senecio nemorensis L. subsp. apuanus (Tausch) Greuter (Asteraceae)

+ EMR: Ventasso (Reggio Emilia), nei pressi del Passo del Cerreto (WGS84: 44.304982°N, 10.223415°E), margine di faggeta lungo strada, 1180 m, 11 August 2025, M. Adorni, C. Zanni (FI); Ventasso (Reggio Emilia), Fiume Secchia a monte de La Gabellina (WGS84: 44.316237°N, 10.224762°E), bosco ripariale di Alnus incana, 1095 m, 11 August 2025 (Herb. M. Adorni) – Subspecies new for the flora of Emilia-Romagna.

Our findings are very close to the populations reported by Marchetti (2019) for the Tuscan side of the Cerreto pass.

M. Adorni, C. Zanni

Thalictrum simplex L. subsp. tenuifolium (Sw. ex Hartm.) Sterner (Ranunculaceae)

+ PIE: Cesana T.se (Torino), zona umida sotto Cappella del Buon Soccorso (WGS84: 44.93189°N, 6.80763°E), 1533 m s.l.m., 28 July 2025, A. Selvaggi (FI, Herb. Selvaggi). – Subspecies confirmed for the flora of Piemonte.

The historical occurrence of this taxon in Piemonte is based on Hand (2001), who cited a specimen preserved at B, collected at “Sestrière, s.d.” by Edoardo Rostan [1826–1895]. The occurrence at Sestrière, geographically close to the new locality documented here, should be considered almost certainly extinct as a consequence of land-use changes that affected the area after 1935 due to the development of the homonymous ski resort.

A. Selvaggi

Trifolium mutabile Port. subsp. gussoneanum (Gibelli & Belli) C.Brullo & Brullo (Fabaceae)

+ BAS: Rionero in Vulture (Potenza), praterie aride lungo la strada fra Rionero in Vulture e i Laghi di Monticchio (WGS84: 40.919903°N, 15.628536°E), 825 m, suolo sciolto, ricco di scheletro, derivante da alterazione di rocce vulcaniche, abbondante, 14 June 2025, G. Salerno, L. Rosati (FI). – Subspecies confirmed for the flora of Basilicata.

In Basilicata, this subspecies was first recorded by Gavioli (1948) as Trifolium vesiculosum Savi var. mutabile Savi. The specimen, dating back to the 1930s and kept in FI! was subsequently revised by Scoppola et al. (2016) as T. mutabile var. gussoneanum.

G. Salerno, L. Rosati

Floristic records of regional alien taxa

Ficus carica L. (Moraceae)

+ (NAT) PIE: Collegno (Torino), cavalcavia di Corso Francia presso via Torino (WGS84: 45.073137°N, 7.587952°E), interstizi del muro di contenimento del cavalcavia, 298 m, 5 June 2025, M. Lonati, G. Nota (FI). – Status change from casual to naturalized alien for the flora of Piemonte.

Ficus carica is a Mediterranean species, considered alien in some northern Italian administrative regions (Bartolucci et al. 2024). In the collection site described here, several plants of different ages and heights colonize a vertical stone wall, forming a stable population. The largest plants are fruit-bearing. The status change from casual to naturalized is consistent with the status documented in the other Italian regions where the species is considered alien.

M. Lonati, G. Nota

Laurus nobilis L. (Lauraceae)

+ (NAT) PIE: Ronco Biellese (Biella), all’interno della Riserva Naturale del Brich di Zumaglia (WGS84: 45.5848°N, 8.0918°E), bosco misto di latifoglie e conifere, 605 m, 17 September 2025, E. Bonivento (FI). – Status change from casual to naturalized alien for the flora of Piemonte.

The species colonizes a mixed broadleaf and conifer forest in the natural reserve “Brich di Zumaglia”, with Pinus strobus L., Picea abies (L.) H.Karst and Castanea sativa Mill. More than one hundred individuals have been found in an area of 2,500 m2, belonging to two different generations. There are mature plants between 6 and 8 m tall as well as younger individuals growing below them. In the undergrowth there is also an abundance of Prunus laurocerasus L.

E. Bonivento

Nerium oleander L. subsp. oleander (Apocynaceae)

+ (NAT) MOL: Campomarino (Campobasso), tra la SS16 Adriatica e la ferrovia (WGS84: 41.934632°N, 15.096622°E) all’interno del sito Natura 2000 ZSC IT7222217 ‘Foce Saccione - Bonifica Ramitelli’, 0 m, E, 26 September 2025, leg. L.A. Santoianni, det. M.C. de Francesco, (FI, IS n. 06313). – Naturalized alien species new for the flora of Molise.

A population of approximately fifty individuals is found in the back-dune zone dominated by Juniperus macrocarpa Sm. and by reforestation plants, such as Pinus halepensis Mill., P. pinaster Aiton and P. pinea L. Furthermore, the presence of other individuals along the Molise coast, both on the more degraded edges of the back-dune areas and in coastal rush beds, suggests that the species is naturalized in this administrative region.

M.C. de Francesco, L.A. Santoianni

Nomenclatural novelties

Anemone alpina L. subsp. austroalpina (D.M.Moser) Galasso & Bartolucci, comb. nov. (Ranunculaceae)

urn:lsid:ipni.org:names:77382078-1

≡ Pulsatilla alpina (L.) Delarbre subsp. austroalpina D.M.Moser, Candollea 58: 46 (2003)

According to recent molecular studies, the genus Anemone L. forms a well-supported monophyletic clade that includes Pulsatilla Mill. as well as the genera Anemonoides Mill., Barneoudia Gay, Eriocapitella Nakai, Hartiana Raf., Knowltonia Salisb., Nemorosa Nieuwl., Miyakea Miyabe & Tatew., Oreithales Schltdl., Oriba Adans., and Preonanthus Ehrh. (Xiang et al. 2026).

F. Bartolucci, G. Galasso

Festuca × stacei (Krahulec) Ardenghi, Banfi & Galasso, comb. nov. (Poaceae)

urn:lsid:ipni.org:names:77382080-1

≡ ×Festulpia stacei Krahulec, Brit. Irish Bot. 8(1): 2 (2026).

A new name, ×Festulpia stacei Krahulec, was recently assigned to the hybrid between Festuca rubra L. and Vulpia myuros (L.) C.C.Gmel. (Krahulec 2026), whose occurrence in Italy was ascertained by Ardenghi et al. (2011). Since phylogenetic treatments currently consider Vulpia C.C.Gmel. as a synonym of Festuca Tourn. ex L. (see for example Soreng et al. 2022), a taxonomic view also shared by Bartolucci et al. (2024), a new combination? for this hybrid under Festuca is provided here.

N. M. G. Ardenghi, E. Banfi, G. Galasso

Nomenclatural and distribution updates from other literature sources

Nomenclatural and distribution updates, and corrigenda to Bartolucci et al. (2024) according to Winkler et al. (2013), Van De Beek (2014, 2016), Kirchmeier et al. (2023), Bertolli et al. (2024), Ferrari (2024a, 2024b), Foelsche (2024), Hanes et al. (2024), Ardenghi (2025a, 2025b), Bovio et al. (2025), Caković et al. (2025), Colli-Silva et al. (2025), Dellavedova et al. (2025), Gariboldi (2025), Giacò et al. (2025), Gottschlich (2025), Gottschlich et al. (2025), Hertel and Presser (2025), Kovalchuk (2025), Nachychko et al. (2025), Pastore and Mota (2025), Perico (2025), Peruzzi (2025, 2026), Prosser et al. (2025), Selvaggi and Jiménez-Mejías (2025), Stinca (2025), Zografidis et al. (2025), Wilhalm and Spögler (2025), Alessandrini et al. (2026), Boucher et al. (2026), Buono et al. (2026), Del Guacchio et al. (2026a, 2026b), Di Gristina et al. (2026), Kadereit and Schneeweiss (2026), Kenny et al. (2026), Lobba et al. (2026), Longo et al. (2026a, 2026b), Manconi and Peruzzi (2026), Manni et al. (2026), Nicolella et al. (2026), Pastore et al. (2026), Roma-Marzio et al. (2026a, 2026b), Samorè et al. (2026), Sukhorukov et al. (2026), Venturi et al. (2026), Verloove et al. (2026), Romero Zarco (2026), Zepigi et al. (2026), Zozomová-Lihová et al. (2026), Xiang et al. (2026) are provided in Suppl. material 1.

F. Bartolucci, G. Galasso

Acknowledgements

We gratefully acknowledge the following colleagues who provided distribution, nomenclatural, and taxonomic advice: Acta Plantarum Staff, Francesco Festi, Duilio Iamonico, Lorenzo Peruzzi, Brunello Pierini. The author (AS) thanks the Ente di Gestione delle Aree Protette delle Alpi Cozie for funding the research within the Interreg VI-A ALCOTRA 2021–2027 project “ACLIMO”.

References

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Additional information

Conflict of interest

The authors have declared that no competing interests exist.

Ethical statement

No ethical statement was reported.

Artificial Intelligence (AI) use

The authors accept full responsibility for the content of the manuscript, including the disclosure of any use of AI.

No AI tools were used in the preparation of this manuscript.

Funding

The contribution of Alberto Selvaggi to this work was funded by the Ente di Gestione delle Aree Protette delle Alpi Cozie within the Interreg VI-A ALCOTRA 2021–2027 project “ACLIMO”.

Author contributions

All authors have contributed equally.

Author ORCIDs

F. Bartolucci https://orcid.org/0000-0001-8199-6003

G. Domina https://orcid.org/0000-0003-4184-398X

A. Alessandrini https://orcid.org/0000-0003-3633-9957

N.M.G. Ardenghi https://orcid.org/0000-0002-4366-5729

G. Bonari https://orcid.org/0000-0002-5574-6067

M.C.a de Francesco https://orcid.org/0000-0002-5238-1154

E. Del Guacchio https://orcid.org/0000-0001-9349-1328

R. Di Pietro https://orcid.org/0000-0003-4983-8931

L. Forte https://orcid.org/0000-0001-8168-7999

G. Galasso https://orcid.org/0000-0002-2501-456X

G. Gottschlich https://orcid.org/0000-0003-0677-6478

M. Lonati https://orcid.org/0000-0001-8886-0328

Q.G. Manni https://orcid.org/0000-0003-4541-3688

F. Minutillo https://orcid.org/0009-0004-4779-591X

G. Nota https://orcid.org/0000-0002-1265-1201

G. Pappagallo https://orcid.org/0000-0002-8565-1130

E. Proietti https://orcid.org/0000-0001-5665-858X

L.A. Santoianni https://orcid.org/0009-0008-3486-0769

A. Selvaggi https://orcid.org/0000-0003-2483-4790

L. Lastrucci https://orcid.org/0000-0003-4455-389X

Data availability

All of the data that support the findings of this study are available in the main text or Supplementary Information.

Supplementary material

Supplementary material 1 

Supplementary data

Fabrizio Bartolucci, Gabriele Galasso

Data type: pdf

Explanation note: 1. Nomenclatural updates; 2. Distribution updates; 3. Synonyms, misapplied or included names.

This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
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