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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">54</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:719A4407-6918-50E6-BCDF-A4165D76DC1E</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">Italian Botanist</journal-title>
        <abbrev-journal-title xml:lang="en">IB</abbrev-journal-title>
      </journal-title-group>
      <issn pub-type="epub">2531-4033</issn>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/italianbotanist.22.212772</article-id>
      <article-id pub-id-type="publisher-id">212772</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Research Article</subject>
        </subj-group>
        <subj-group subj-group-type="biological_taxon">
          <subject>Asteraceae</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Nomenclature</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>Typification of the Linnaean name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family" reg="Asteraceae">Asteraceae</tp:taxon-name-part></tp:taxon-name>)</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Abdollahi</surname>
            <given-names>Mohammad Reza</given-names>
          </name>
          <email xlink:type="simple">mo.abdollahi5@gmail.com</email>
          <uri content-type="orcid">https://orcid.org/0009-0002-8411-0495</uri>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Peruzzi</surname>
            <given-names>Lorenzo</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-9008-273X</uri>
          <xref ref-type="aff" rid="A2">2</xref>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Ghahremaninejad</surname>
            <given-names>Farrokh</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-5860-9976</uri>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">Department of Plant Sciences, Faculty of Biological Sciences, Kharazmi University, Postal Code 15719–14911, Tehran, Iran</addr-line>
        <institution>Department of Plant Sciences, Faculty of Biological Sciences, Kharazmi University</institution>
        <addr-line content-type="city">Tehran</addr-line>
        <country>Iran</country>
        <uri content-type="ror">https://ror.org/05hsgex59</uri>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">Department of Biology, PLANTSEED Lab, University of Pisa, Via Derna 1, 56126 Pisa, Italy</addr-line>
        <institution>Department of Biology, PLANTSEED Lab, University of Pisa</institution>
        <addr-line content-type="city">Pisa</addr-line>
        <country>Italy</country>
        <uri content-type="ror">https://ror.org/03ad39j10</uri>
      </aff>
      <aff id="A3">
        <label>3</label>
        <addr-line content-type="verbatim">Botanic Garden and Museum, University Museum System, University of Pisa, Via Luca Ghini 13, 56126 Pisa, Italy</addr-line>
        <institution>Botanic Garden and Museum, University Museum System, University of Pisa</institution>
        <addr-line content-type="city">Pisa</addr-line>
        <country>Italy</country>
        <uri content-type="ror">https://ror.org/03ad39j10</uri>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Mohammad Reza Abdollahi (<ext-link xlink:href="mailto:mohammad.abdollahi@khu.ac.ir" ext-link-type="uri">mohammad.abdollahi@khu.ac.ir</ext-link>)</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: Simone Orsenigo</p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2026</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>21</day>
        <month>09</month>
        <year>2026</year>
      </pub-date>
      <volume>22</volume>
      <fpage>65</fpage>
      <lpage>70</lpage>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/094E76BE-B52E-52CA-87FF-D3D3B6492098">094E76BE-B52E-52CA-87FF-D3D3B6492098</uri>
      <history>
        <date date-type="received">
          <day>23</day>
          <month>08</month>
          <year>2026</year>
        </date>
        <date date-type="accepted">
          <day>11</day>
          <month>09</month>
          <year>2026</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Mohammad Reza Abdollahi, Lorenzo Peruzzi, Farrokh Ghahremaninejad</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <p>The typification of the Linnaean name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family" reg="Asteraceae">Asteraceae</tp:taxon-name-part></tp:taxon-name>) is discussed. Examination of the protologue and associated original material revealed that both the illustration published by Gmelin in “Flora Sibirica” and a specimen preserved in the Linnaean Herbarium (Herb. Linn. No. 1017.9, <abbrev xlink:title="Linnaean Herbarium">LINN</abbrev>) constitute original material for the name. To promote nomenclatural stability and preserve current usage, Herb. Linn. No. 1017.9 (<abbrev xlink:title="Linnaean Herbarium">LINN</abbrev>) is here designated as the lectotype of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">A.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>.</p>
      </abstract>
      <kwd-group>
        <label>Keywords</label>
        <kwd>
          <italic>
            <tp:taxon-name>
              <tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part>
            </tp:taxon-name>
          </italic>
        </kwd>
        <kwd>
          <tp:taxon-name>
            <tp:taxon-name-part taxon-name-part-type="family" reg="Compositae">Compositae</tp:taxon-name-part>
          </tp:taxon-name>
        </kwd>
        <kwd>Linnaean name</kwd>
        <kwd>nomenclature</kwd>
        <kwd>typification</kwd>
      </kwd-group>
    </article-meta>
    <notes>
      <sec sec-type="Citation" id="sec1">
        <title>Citation</title>
        <p>Abdollahi MR, Peruzzi L, Ghahremaninejad F (2026) Typification of the Linnaean name <italic>Achillea impatiens</italic> (Asteraceae). Italian Botanist 22: 65–70. <ext-link xlink:href="10.3897/italianbotanist.22.212772" ext-link-type="doi">https://doi.org/10.3897/italianbotanist.22.212772</ext-link></p>
      </sec>
    </notes>
  </front>
  <body>
    <sec sec-type="Introduction" id="sec2">
      <title>Introduction</title>
      <p>The genus <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part></tp:taxon-name></italic> L. (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family" reg="Asteraceae">Asteraceae</tp:taxon-name-part></tp:taxon-name> Bercht. &amp; J.Presl), commonly known as Yarrow, is one of the largest genera within the tribe <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="tribe" reg="Anthemideae">Anthemideae</tp:taxon-name-part></tp:taxon-name> Cass., currently comprising approximately 141 accepted species distributed throughout the temperate and subtropical regions of the Northern Hemisphere, with centres of diversity in southeastern Europe and southwestern Asia and a distribution extending across Eurasia to North America (<xref ref-type="bibr" rid="B2">Bremer and Humphries 1993</xref>; <xref ref-type="bibr" rid="B8">Guo et al. 2004</xref>; <xref ref-type="bibr" rid="B4">Ehrendorfer and Guo 2006</xref>; <xref ref-type="bibr" rid="B5">Euro+Med 2026</xref>; <xref ref-type="bibr" rid="B15">POWO 2026</xref>).</p>
      <p>Carl Linnaeus (<xref ref-type="bibr" rid="B12">1753</xref>, <xref ref-type="bibr" rid="B13">1759</xref>, <xref ref-type="bibr" rid="B14">1763</xref>) published 22 names in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part></tp:taxon-name></italic> (<xref ref-type="bibr" rid="B10">Jarvis 2007</xref>; <xref ref-type="bibr" rid="B22">Wajer 2022</xref>; IPNI 2026). Most of these names are currently accepted, whereas five are now treated as synonyms following subsequent taxonomic revisions, including transfers to other genera or changes in taxonomic rank [<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="bipinnata">bipinnata</tp:taxon-name-part></tp:taxon-name></italic> L. = <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Tanacetum">Tanacetum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="argyrophyllum">argyrophyllum</tp:taxon-name-part></tp:taxon-name></italic> (K.Koch) Tzvelev; <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="inodora">inodora</tp:taxon-name-part></tp:taxon-name></italic> L. = <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Lonas">Lonas</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="annua">annua</tp:taxon-name-part></tp:taxon-name></italic> Vines &amp; Druce; <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="magna">magna</tp:taxon-name-part></tp:taxon-name></italic> L. = <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="millefolium">millefolium</tp:taxon-name-part></tp:taxon-name></italic> subsp. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea"/><tp:taxon-name-part taxon-name-part-type="species">millefolium</tp:taxon-name-part></tp:taxon-name>; <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="santolina">santolina</tp:taxon-name-part></tp:taxon-name></italic> L. = <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="tenuifolia">tenuifolia</tp:taxon-name-part></tp:taxon-name></italic> Lam.; <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="abrotanifolia">abrotanifolia</tp:taxon-name-part></tp:taxon-name></italic> L. = <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Tanacetum">Tanacetum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="abrotanifolium">abrotanifolium</tp:taxon-name-part></tp:taxon-name></italic> (L.) Druce)] (<xref ref-type="bibr" rid="B22">Wajer 2022</xref>). All the accepted species described by Linnaeus have been typified through previous nomenclatural studies (<xref ref-type="bibr" rid="B6">Franzén 1986</xref>; <xref ref-type="bibr" rid="B19">Valant-Vetschera 1995</xref>; <xref ref-type="bibr" rid="B11">Jarvis and Turland 1998</xref>; <xref ref-type="bibr" rid="B10">Jarvis 2007</xref>), with the exception of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> that remains untypified (<xref ref-type="bibr" rid="B22">Wajer 2022</xref>). The present study aims to stabilize the application of this Linnaean name through lectotypification.</p>
    </sec>
    <sec sec-type="materials|methods" id="sec3">
      <title>Materials and methods</title>
      <p>This study is based on the analysis of the protologue of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> published by Linnaeus (1753: 898) and the works cited therein, particularly those by van Royen (1740) and <xref ref-type="bibr" rid="B7">Gmelin (1747)</xref>. Additional historical sources relevant to the interpretation of the name, including <xref ref-type="bibr" rid="B3">Candolle (1838)</xref>, were also examined. High-resolution digital images of specimens preserved in the Linnaean Herbarium (<abbrev xlink:title="Linnaean Herbarium">LINN</abbrev>) and the Smith Herbarium (<abbrev xlink:title="Smith Herbarium">LINN-HS</abbrev>) were studied through the Linnean Collections Online portal (<ext-link xlink:href="https://linnean.access.preservica.com/" ext-link-type="uri">https://linnean.access.preservica.com/</ext-link>). Original illustrations published in “Flora Sibirica” (<xref ref-type="bibr" rid="B7">Gmelin 1747</xref>) were consulted using digital scans available through the digital library of the Real Jardín Botánico de Madrid (<ext-link xlink:href="https://bibdigital.rjb.csic.es/" ext-link-type="uri">https://bibdigital.rjb.csic.es/</ext-link>). Typification follows the rules of the “International Code of Nomenclature for algae, fungi, and plants” (<xref ref-type="bibr" rid="B18">Turland et al. 2025</xref>; <abbrev xlink:title="International Code of Nomenclature">ICN</abbrev> hereafter).</p>
    </sec>
    <sec sec-type="Typification of the name" id="sec4">
      <title>Typification of the name</title>
      <tp:taxon-treatment>
        <tp:treatment-meta>
          <kwd-group>
            <label>Taxon classification</label>
            <kwd>
              <named-content content-type="kingdom">Animalia</named-content>
            </kwd>
            <kwd>
              <named-content content-type="order">Asterales</named-content>
            </kwd>
            <kwd>
              <named-content content-type="family">Asteraceae</named-content>
            </kwd>
          </kwd-group>
        </tp:treatment-meta>
        <tp:nomenclature>
          <tp:taxon-name><object-id content-type="arpha">3C541023-502F-58F1-9630-717491F6ED62</object-id>
                		<tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part>
                	</tp:taxon-name>
          <tp:taxon-authority>L., Sp. Pl. 2: 898. 1753</tp:taxon-authority>
          <tp:nomenclature-citation-list>
            <tp:nomenclature-citation>
              <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica"/> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens"/></tp:taxon-name>
              <comment> ≡ <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica">Ptarmica</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name> (L.) DC., Prodr. 6: 22. 1838.</comment>
            </tp:nomenclature-citation>
          </tp:nomenclature-citation-list>
        </tp:nomenclature>
        <tp:treatment-sec sec-type="Lectotypus">
          <title>Lectotypus</title>
          <p><bold>(here designated)</bold>. Herb. Linn. No. 1017.9 (<abbrev xlink:title="Linnaean Herbarium">LINN</abbrev> [digital image!]; Fig. <xref ref-type="fig" rid="F1">1</xref>).</p>
          <fig id="F1">
            <object-id content-type="doi">10.3897/italianbotanist.22.212772.figure1</object-id>
            <object-id content-type="arpha">5051C146-A26A-5905-B5D2-737A68E52E24</object-id>
            <label>Figure 1.</label>
            <caption>
              <p>Lectotype of the name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> L. (Herb. Linnaeus No. 1017.9 in <abbrev xlink:title="Linnaean Herbarium">LINN</abbrev>) (Reproduced with permission under a CC BY-NC 4.0 license).</p>
            </caption>
            <graphic xlink:href="italianbotanist-22-065-g001.jpg" id="oo_1785640.jpg">
              <uri content-type="original_file">https://binary.pensoft.net/fig/1785640</uri>
            </graphic>
          </fig>
        </tp:treatment-sec>
        <tp:treatment-sec sec-type="Remarks">
          <title>Remarks.</title>
          <p>The name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> was described by Linnaeus (1753: 898) as “8. ACHILLEA foliis pinnatis: pinnis distantibus lineari-lanceolatis: basi sursum auctis”; this diagnosis was taken directly from van Royen (1740: 176); a polynomial published by Gmelin (1747: 197, t. 83, f. 1; “Achillea foliis pinnatis, pinnis longis acutis: inferioribus sursum dente auctis glaberrimis”) was also cited; the provenance was given as “Habitat in Sibiria”. In addition, a description of the plant (“Planta albis gaudet floribus. Foliorum pinnae remotae et admodum distantes, distinguunt plantam a reliquis”) was reported.</p>
          <p>Among the elements associated with the protologue of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>, the illustration published by Gmelin in “Flora Sibirica” (1747: 197, tab. LXXXIII, fig. I) (see <ext-link xlink:href="https://bibdigital.rjb.csic.es/viewer/12378/?offset=#page=357&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" ext-link-type="uri">https://bibdigital.rjb.csic.es/viewer/12378/?offset=#page=357&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=</ext-link>) constitutes original material because it was explicitly cited by Linnaeus in the protologue. A specimen preserved in the Linnaean Herbarium at <abbrev xlink:title="Linnaean Herbarium">LINN</abbrev> (Herb. Linn. No. 1017.9; numbering follows <xref ref-type="bibr" rid="B17">Savage 1945</xref>) has been also traced. The sheet bears the Linnaean annotation “8 impatiens”, where the number “8” corresponds to the Linnaean number of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">A.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> in “Species Plantarum”. This means that <abbrev xlink:title="Linnaean Herbarium">LINN</abbrev>-1017.9 is part of the original material for the name. The specimen is well preserved and displays diagnostic features consistent with both the protologue and Gmelin’s illustration. At <abbrev xlink:title="Smith Herbarium">LINN-HS</abbrev> (Herb. Smith), two further specimens are preserved on sheet No. 1315.24 (Nos. 1315.24.1 and 1315.24.2; image available at <ext-link xlink:href="https://linnean.access.preservica.com/uncategorized/IO_f317aecb-4592-4cf3-ad3a-c32031611710/" ext-link-type="uri">https://linnean.access.preservica.com/uncategorized/IO_f317aecb-4592-4cf3-ad3a-c32031611710/</ext-link>). The original annotations on the sheet provide explicit evidence of their post-1753 provenance. Specimen 1 is labelled “Rev. Mr. Watts’s garden, Ashill. Sep. 1. 1801”, while specimen 2 is labelled “From his herbarium”; the sheet also bears the later annotations “Achillea impatiens” and “Mr Watts”. Thus, specimen 1 was obtained from Rev. Mr. Watts’s garden on 1 September 1801, 48 years after the publication of “Species Plantarum” (1753), while specimen 2 is explicitly identified as having come from his herbarium. The explicit post-1753 provenance of the specimens demonstrates that they were not available to Linnaeus when he prepared the protologue of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">A.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>. Accordingly these specimens cannot be considered original material for the name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">A.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>.</p>
          <p>No specimen linked to the diagnosis cited by Linnaeus from van Royen (1740) could be located in the van Royen Herbarium (preserved at L).</p>
          <p>All original elements identified are consistent with the protologue of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>. Both the specimen at <abbrev xlink:title="Linnaean Herbarium">LINN</abbrev> (No. 1017.9) and the illustration published by Gmelin (“Flora Sibirica” 2: 197, tab. LXXXIII, fig. 1) exhibit the diagnostic characters emphasized by Linnaeus, including pinnately divided leaves with elongate, acute, linear-lanceolate segments, the lowermost segments bearing an additional upward-directed tooth, and the distinctly remote arrangement of the leaf segments, leaving the rachis conspicuously exposed, a feature that Linnaeus explicitly regarded as distinguishing the species from the remaining species of the genus.</p>
          <p>In accordance with Art. 9.12 of the <abbrev xlink:title="International Code of Nomenclature">ICN</abbrev>, the specimen Herb. Linn. No. 1017.9 (<abbrev xlink:title="Linnaean Herbarium">LINN</abbrev>) (Fig. <xref ref-type="fig" rid="F1">1</xref>) is here designated as the lectotype of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>. The lectotype matches the protologue and the traditional concept and current application of the name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">A.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> (e.g., <xref ref-type="bibr" rid="B9">Heimerl 1884</xref>; <xref ref-type="bibr" rid="B1">Afanasyev and Bochantsev 1961</xref>; <xref ref-type="bibr" rid="B16">Richardson 1976</xref>; <xref ref-type="bibr" rid="B8">Guo et al. 2004</xref>; <xref ref-type="bibr" rid="B4">Ehrendorfer and Guo 2006</xref>).</p>
          <p>Candolle (<xref ref-type="bibr" rid="B3">1838</xref>: 22) accepted this species under the genus <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica">Ptarmica</tp:taxon-name-part></tp:taxon-name></italic> Mill. and published the name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica">Ptarmica</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic>. In the protologue, he explicitly cited the name <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> L., as well as the main synonyms associated with the Linnaean concept of this species, including Gmelin’s illustration. Consequently, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica">Ptarmica</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> is a nomenclatural combination based on <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Achillea">Achillea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> L., i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica">Ptarmica</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> (L.) DC. (Art. 7.3 of the <abbrev xlink:title="International Code of Nomenclature">ICN</abbrev>), although <xref ref-type="bibr" rid="B15">POWO (2026)</xref> erroneously treats this name as a heterotypic synonym (i.e. <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ptarmica">Ptarmica</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="impatiens">impatiens</tp:taxon-name-part></tp:taxon-name></italic> “DC.”).</p>
        </tp:treatment-sec>
      </tp:taxon-treatment>
    </sec>
  </body>
  <back>
    <ack>
      <title>Acknowledgements</title>
      <p>The authors gratefully acknowledge the Linnean Society of London for providing online access to the Linnaean Collections. The authors also thank M. Hjertson (Uppsala) for his helpful advice on the nomenclatural aspects of this study.</p>
    </ack>
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    <sec sec-type="Additional information" id="sec5">
      <title>Additional information</title>
      <sec sec-type="Conflict of interest" id="sec6">
        <title>Conflict of interest</title>
        <p>The authors have declared that no competing interests exist.</p>
      </sec>
      <sec sec-type="Ethical statement" id="sec7">
        <title>Ethical statement</title>
        <p>No ethical statement was reported.</p>
      </sec>
      <sec sec-type="Artificial Intelligence (AI) use" id="sec8">
        <title>Artificial Intelligence (AI) use</title>
        <p>The authors accept full responsibility for the content of the manuscript, including the disclosure of any use of AI.</p>
        <p>No AI tools were used in the preparation of this manuscript.</p>
      </sec>
      <sec sec-type="Funding" id="sec9">
        <title>Funding</title>
        <p>No funding was reported.</p>
      </sec>
      <sec sec-type="Author contributions" id="sec10">
        <title>Author contributions</title>
        <p>All authors have contributed equally.</p>
      </sec>
      <sec sec-type="Author ORCIDs" id="sec11">
        <title>Author ORCIDs</title>
        <p>M.R. Abdollahi <ext-link xlink:href="https://orcid.org/0009-0002-8411-0495" ext-link-type="uri">https://orcid.org/0009-0002-8411-0495</ext-link></p>
        <p>L. Peruzzi <ext-link xlink:href="https://orcid.org/0000-0001-9008-273X" ext-link-type="uri">https://orcid.org/0000-0001-9008-273X</ext-link></p>
        <p>F. Ghahremaninejad <ext-link xlink:href="https://orcid.org/0000-0001-5860-9976" ext-link-type="uri">https://orcid.org/0000-0001-5860-9976</ext-link></p>
      </sec>
      <sec sec-type="Data availability" id="sec12">
        <title>Data availability</title>
        <p>All of the data that support the findings of this study are available in the main text.</p>
      </sec>
    </sec>
  </back>
</article>
